British Wild Flowers: A photographic guide to every common species
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Collins
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First published in 2006
Text © 2006 Paul Sterry
Photographs © Individual photographers indicated in the picture credits
Colour reproduction by Nature Photographers Ltd.
Paul Sterry asserts his moral right to be identified as the author of this work
A catalogue record for this book is available from the British Library
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Source ISBN: 9780007236848
Ebook Edition © MARCH 2015 ISBN: 9780008144586
Version: 2016-12-05
CONTENTS
Cover
Title Page
Copyright
Introduction
How to Use this Book
Basic Botany
Glossary
Flowers
Wild Flower Identification
Fruits and Seeds
Leaves
Habitats
Conservation
Species Descriptions
Juniper to Nettle Families
Nettle, Birthwort and Knotweed Families
Knotweed Family Polygonaceae
Blinks to Goosefoot Families
Goosefoot Family Chenopodiaceae
Goosefoot and Pink Families
Pink Family Caryophyllaceae
Buttercup Family Ranunculaceae
Fumitory and Poppy Families
Poppy, Barberry and Cabbage Families
Cabbage Family Brassicaceae
Cabbage and Mignonette Families
Sundew, Pitcherplant and Stonecrop Families
Stonecrop and Saxifrage Families
Saxifrage Family Saxifrageaceae
Gooseberry and Rose Families
Rose Family Rosaceae
Pea Family Fabaceae
Pea and Wood-sorrel Families
Crane’s-bill Family Geraniaceae
Crane’s-bill and Flax Families
Spurge Family Euphorbiaceae
Spurge, Milkwort and Balsam Families
Holly, Spindle, Buckthorn, Box and Mallow Families
Mallow to St John’s-wort Families
St John’s-wort and Rock-rose Families
Violet Family Violaceae
Violet to Willowherb Families
Waterwort and Willowherb Families
Willowherb and Dogwood Families
Ivy and Carrot Families
Carrot Family Apiaceae
Primrose Family Primulaceae
Heather Family Ericaceae
Heather and Crowberry Families
Wintergreen, Bird’s-nest and Thrift Families
Ash and Gentian Families
Gentian, Bogbean, Periwinkle and Bedstraw Families
Bedstraw Family Rubiaceae
Bedstraw, Jacob’s-ladder and Dodder Families
Bindweed and Borage Families
Borage Family Boraginaceae
Vervain and Dead-nettle Families
Dead-nettle Family Lamiaceae
Dead-nettle and Butterfly-bush Families
Nightshade and Figwort Families
Figwort Family Scrophulariaceae
Figwort and Broomrape Families
Broomrape, Moschatel and Valerian Families
Valerian and Butterwort Families
Plantain and Arrowgrass Families
Honeysuckle and Teasel Families
Teasel and Bellflower Families
Bellflower Family Campanulaceae
Daisy Family Asteraceae
Iris, Black Bryony and Lords-and-ladies Families
Orchid Family Orchidaceae
Water Plants
Pondweeds Potamogetonaceae
Bur-reed, Bulrush and Rush Families
Rush and Sedge Families
Sedge Family Cyperaceae
Sedge and Grass Families
Grass Family Poaceae
The West Country
The Lizard
The Isles of Scilly
Channel Islands
The New Forest and Isle of Wight
South-East England’S Estuaries and Coasts
The South and North Downs
East Anglia
The Chilterns and Cotswolds
English and Welsh Uplands
Scottish Highlands and Western Isles
Shetland and Orkney Isles
Ireland
Further Reading and Useful Organisations
Index
About the Publisher
INTRODUCTION
PEOPLE LIVING IN BRITAIN and Ireland seldom have to travel far to find a wealth of wild flowers and, although the region’sv plantlife faces significant conservation issues, residents should feel privileged to live in such a flower-rich part of the world. Underpinning our floral diversity is a rich array of habitats, the product of our region’s topography, geology and history of land use. Good fortune, in the form of the Gulf Stream, dictates a mild and comparatively equable climate for much of the time, and this, too, contributes to botanical diversity. Complete British Wild Flowers has arisen from my personal love for the flora of Britain and Ireland, not to mention half a lifetime devoted to photographing our wild flowers.
THE REGION COVERED BY THIS BOOK
The region covered by this book comprises the whole of mainland England, Wales, Scotland and Ireland, as well as offshore islands including the Shetlands, Orkneys, Hebrides, Isle of Man and the Isles of Scilly. In addition, I have included the Channel Islands because their proximity to, and ecological affinities with, northern France allow them to make a valuable contribution to our flora.
THE CHOICE OF SPECIES
The coverage of the book is restricted mainly to what most people understand to be wild flowers – not a strict botanical term, but taken to mean flowering plants of relatively modest stature, species that in most cases do not exceed 2m in height. For the sake of completeness I have also included a number of woody flowering shrubs, but I have excluded obvious tree species. Terrestrial habitats harbour the lion’s share of our flowering plants and this is reflected in the coverage of this book. But I have also included species that are strictly aquatic, both those that occur in freshwater habitats and the limited range of flowering plants that grow in coastal marine environments.
Complete British Wild Flowers is aimed primarily at the botanical novice and those with a moderate degree of botanical experience. Consequently, for the main section of the book, I have selected species that the naturalist stands a reasonable chance of encountering, although a few scarce but spectacular and distinctive species have also been included here for good measure. Some botanical groups are minefields for the beginner: for example, hundreds of so-called ‘microspecies’ of brambles are recognised by experts and similar complexity is found among dandelions, hawkweeds and eyebrights to name three more. In a book of this size, accurate identification of these subdivisions is impossible to achieve. So rather than baffle the reader, I have chosen to ‘lump’ the subdivisions together within these complex groups and treat them as species aggregates.
In the hope and expectation that readers’ appetites for botanical discoveries are whetted by the book, the last section of Complete British Wild Flowers is devoted to botanical hotspots, areas that merit a visit in search of the localised floral treasures they harbour.
Floral delights such as this heathland have inspired generations of botanists.
HOW TO USE THIS BOOK
THE BOOK HAS BEEN designed so that the text and photographs for each species are on facing pages. A system of labelling clearly states the flower’s identity. The text has been written to complement the information conveyed by the photographs. By and large, the order in which the species appear in the main section of the book roughly follows standard botanical classification. However, because parts of the field are in a state of flux, the order may differ slightly from that found in other guides, past, present or future.
SPECIES DESCRIPTIONS
At the start of each species description the plant’s most commonly used and current English name is given. This is followed by the scientific name, comprising the genus name first and then by the specific name. In a few instances, where this is pertinent, reference is made, in either the species heading or the main body of the text, to a further subdivision: subspecies. In general, I have followed the nomenclature of Clive Stace’s New Flora of the British Isles (see p.). However, with orchids I have used the most up-to-date classification system, which differs from that in most currently available floras. The family group to which the plant in question belongs is then given. If a species is anything other than widespread and common, there then follows a measure of its scarcity, indicated by rosette symbols:
These scarcity ratings are based on my own observations, so they are subjective and are not intended as an absolute guide to a species’ status. Lastly in the heading, some measure is given of the plant’s size. Size and stature vary according to a number of factors so these values should be treated with a degree of caution.
The text has been written in as concise a manner as possible. Each description begins with a summary of the plant and the habitat that it favours. To avoid potential ambiguities, the following headings break up the rest of each description: FLOWERS; FRUITS; LEAVES; STATUS. These sections describe the colour, shape and size of the various plant parts – features that tend to be more constant, and hence more useful for identification, than the overall size of the plant. Text in italics relates to key features of identification.
MAPS
The maps provide invaluable information about the distribution and occurrence of each species in the region: the intensity of the colour gives an indication of abundance. In compiling the maps, I have made reference to a number of sources, including An Atlas of the Wild Flowers of Britain and Northern Europe, the New Atlas of the British and Irish Flora (see p.), various county floras and, of course, my own notes. The maps represent the current ranges of plants in the region in general terms. Please bear in mind that, given the size of the maps, small and isolated populations will not necessarily be featured. Note also that the ranges of many species (particularly those influenced by agriculture) are likely to contract further as the years go by.
PHOTOGRAPHS
Great care has gone into the selection of photographs for this book and in many cases the images have been taken specifically for this project. Preference has been given to photographs that serve both to illustrate key identification features of a species when in flower, and to emphasise its beauty. In many instances, smaller inset photographs illustrate useful identification features that are not shown clearly by the main image.
BASIC BOTANY
IN COMMON WITH OTHER living organisms, plants have the abilities to grow, reproduce themselves, respond to stimuli and – to a limited extent compared to most animals – move. What sets them apart from animals in particular, and defines plants as a group, is their ability, in almost all species, to manufacture their own food from inorganic building blocks. Fundamental to this ability is the process called photosynthesis, in which plants use energy from sunlight to convert carbon dioxide from the air and water into glucose. Oxygen is a by-product of this reaction. A pigment called chlorophyll is the agent that extracts from sunlight the energy needed to fuel the reaction; its coloration is what makes plants green.
The plants covered in this book are the most advanced of their kind in evolutionary terms. All have the ability to reproduce sexually and flowers are the means by which this process is achieved. Flower structure is as varied as it is complex, and the fact that whole books, including this one, can be devoted to detailing its diversity is testament to this. Flower structure is dealt with in more detail on page 10.
Among terrestrial plants the process of photosynthesis is difficult to demonstrate outside the laboratory. However, certain submerged aquatic plants, such as this Water Starwort, produce bubbles of oxygen when they are exposed to bright sunlight, indicating that photosynthesis is indeed taking place.
THE ROLE OF PLANTS IN ECOLOGY
The importance of plants in the global ecosystem, and at the local level, cannot be overstated. Not only do plants generate (through photosynthesis) the atmospheric oxygen that all animals need to breathe but, as a source of food, they underpin food chains across the world. Without plants, life on earth as we know it would not have evolved and could not survive.
The survival of plants and animals is inextricably linked at the general level, but there are plenty of highly specific instances of dependence, the relationship often hinted at by their English names. Without their namesake-specific foodplants, neither the Figwort Weevil (left) nor the Mullein Moth (right), shown here as a caterpillar, could survive.
GLOSSARY
Achene – one-seeded dry fruit that does not split.
Acute – sharply pointed.
Alien – introduced by man from another part of the world.
Alternate – not opposite.
Annual – plant that completes its life cycle within 12 months.
Anther – pollen-bearing tip of the stamen.
Appressed (sometimes written as adpressed in other books) – pressed closely to the relevant part of the plant.
Auricle – pair of lobes at the base of a leaf.
Awn – stiff, bristle-like projection, seen mainly in grass flowers.
Axil – angle between the upper surface or stalk of a leaf and the stem on which it is carried.
Basal – appearing at the base of plant, at ground level.
Basic – soil that is rich in alkaline (mainly calcium) salts.
Beak – elongate projection at the tip of a fruit.
Berry – fleshy, soft-coated fruit containing several seeds.
Biennial – plant that takes two years to complete its life cycle.
Bog – wetland on acid soil.
Bract – modified, often scale-like, leaf found at the base of flower stalks in some species.
Bracteole – modified, often scale-like, leaf found at the base of individual flowers in some species.
Bulb – swollen underground structure containing the origins of the following year’s leaves and buds.
Bulbil – small, bulb-like structure, produced asexually by some plants and capable of growing into a new plant.
Calcareous – containing calcium, the source typically being chalk or limestone.
Calyx – outer part of a flower, comprising the sepals.
Capsule – dry fruit that splits to liberate its seeds.
Catkin – hanging spike of tiny flowers.
Chlorophyll – green pigment, present in plant leaves and other structures, and essential in the process of photosynthesis.
Cladode – green, leaf-like shoot.
Clasping – referring to leaf bases that have backward-pointing lobes that wrap around the stem.
Composite – member of the daisy family (Asteraceae).
Compound – leaf that is divided into a number of leaflets.
Cordate – heart-shaped at the base.
Corm – swollen underground stem.
Corolla – collective term for the petals.
Cultivar – plant variety created by cultivation.
Deciduous – plant whose leaves fall in autumn.
Decurrent – with the leaf base running down the stem.
Dentate – toothed.
Digitate – resembling the fingers of a splayed hand.
Dioecious – having male and female flowers on separate plants.
Disc floret – one of the inner florets of a composite flower.
Drupe – succulent fruit, the seed inside having a hard coat.
Emergent – a plant growing with its base and roots in water, the rest of the plant emerging above water level.
Entire – in the context of a leaf, a margin that is untoothed.
Epicalyx – calyx-like structure, usually surrounding, and appressed, to the calyx.
Epiphyte – plant that grows on another plant, on which it is not a parasite.
Fen – wetland habitat on alkaline peat.
Filament – stalk part of a stamen.
Flexuous – wavy.
Floret – small flower, part of larger floral arrangement as in composite flowers or umbellifers.
Fruits – seeds of a plant and their associated structures.
Genus (plural Genera) – group of closely related species, sharing the same genus name.
Glabrous – lacking hairs.
Gland – sticky structure at the end of a hair.
Glaucous – blue-grey in colour.
Globose – spherical or globular.
Glume – pair of chaff-like scales at the base of a grass spikelet.
Hybrid – plant derived from the crossfertilisation of two different species.
Inflorescence – the flowering structure in its entirety, including bracts.
Introduced – not native to the region.
Keel – seen in pea family members; the fused two lower petals that are shaped like a boat’s keel.
Lanceolate – narrow and lance-shaped.
Latex – milky fluid.
Lax – open, not dense.
Leaflet – leaf-like segment or lobe of a leaf.
Ligule – somewhat membranous flap at the base of a grass leaf, where it joins the stem.
Linear – slender and parallel-sided.
Lip – usually the lower part of an irregular flower such as an orchid.
Lobe – division of a leaf.
Microspecies – division within a species, members of which are only subtly different from members of other microspecies.
Midrib – central vein of a leaf.
Native – occurring naturally in the region and not known to have been introduced.
Node – point on the stem where a leaf arises.
Nut – a dry, one-seeded fruit with a hard outer case.
Nutlet – small nut.
Oblong – leaf whose sides are at least partly parallel-sided.
Obtuse – blunt-tipped (usually in the context of a leaf).
Opposite – (usually leaves) arising in opposite pairs on the stem.
Opposite
Oval – leaf shape. Ovate is oval in outline.
Ovary – structure containing the ovules, or immature seeds.
Ovoid – egg-shaped.
Palmate – leaf with finger-like lobes arising from the same point.
Panicle – branched inflorescence.
Pappus – tuft of hairs on a fruit.
Parasite – plant that derives its nutrition entirely from another living organism.
Pedicel – stalk of an individual flower.